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  • TITLE
  • DEDICATION
  • DECLARATION
  • CERTIFICATE
  • ACKNOWLEDGEMENT
  • CONTENTS
  • ABBREVIATIONS
  • List of tables
  • LIST OF FIGURES
  • 1. Introduction
  • 1.1 Rubber tree -- An eco-friendly source of timber
  • 1.2 Availability of rubber wood
  • 1.3 Commercial utilization of rubber wood
  • 1.3.1 The Indian scenario
  • 1.4 Demerits of rubber wood
  • 1.5 Relevance of the study
  • 2. Review of literature
  • 2.1 Structural features of rubber wood
  • 2.1.1 Gross structure
  • 2.1.2 Anatomy of rubber wood
  • 2.2 Major demerits of rubber wood
  • 2.2.1 Susceptibility to biological deterioration
  • 2.2.2 Tension wood -- a natural defect
  • 2.3 Structure of tension wood fibres
  • 2.4 Mechanism of tension wood formation
  • 2.5 Tension wood formation in Hevea brasiliensis
  • 2.6 Tension wood - wood working problems
  • 3. Materials and methods
  • 3.1 Distribution pattern, proportion and structure of tension wood in mature trees
  • 3.1.1 Materials
  • 3.1.2 Methodology
  • 3.1.2.1 Collection of wood discs
  • 3.1.2.2 Preparation of samples
  • 3.1.2.3 Quantification of tension wood
  • 3.2 Effect of leaning angle on tension wood formation in mature trees
  • 3.3 Distribution pattern and proportion of tension wood in immature plants
  • 3.4 Tension wood formation in bud grafted and non -bud -grafted (tissue culture) plants
  • 3.5 Tension wood formation in tapped and untapped zones of the trunk
  • 3.6 Tension wood formation and wind damage
  • 3.7 Structural studies on tension wood
  • 3.7.1 Dimension of fibres and vessel elements
  • 3.7.2 Fibre wall thickness
  • 3.7.3 Analysis of pores
  • 3.7.4 Analysis of rays
  • 3.8 Histochemical studies
  • 3.9 Visual identification of tension wood in rubber wood logs
  • 3.10 Factors affecting tension wood formation
  • 3.10.1 Angle of leaning
  • 3.10.2 Angle of leaning with defoliation
  • 3.10.3 Effect of gravity on tension wood formation
  • 3.10.4 Effect of growth regulators
  • 3.10.4.1 Apical application of IAA, TIBA and GA3 on artificially bent plants
  • 3.10.4.2 Lateral application of IAA, TIBA and GA3 on plants grown vertical
  • 3.10.4.3 Lateral application of IAA on plants grown vertical through incision of bark
  • 3.10.4.4 Lateral application of IAA and TIBA on plants under artificial bending
  • 3.11 Statistical Analysis
  • 3.12 Photography
  • 1 Diagrammatic representation of collection wood discs from tree and preparation of samples
  • 2 Tissue culture plant and budgrafted plant at the age of 8 months (clone R.RII 105)
  • 3 Angle of leaning seedlings bent at 45° and 90°
  • 4 Angle of leaning: seedling.bent at 135°
  • 5 Angle of leaning with defoliation: seedlings bent at 45° and 90°
  • 6 Angle of leaning with defoliation: seedling bent at 135° 7 Loop Experiment
  • 7. Loop Experiment
  • 8 Method of application of growth regulators on bent and vertically growing plants
  • 9 Method of application of growth regulators on bent and vertically growing plants
  • 4. Results
  • 4.1 Distribution pattern and proportion of tension wood in mature trees
  • 4.2 Distribution pattern and proportion of tension wood in immature plants
  • 4.3 Tension wood formation in bud-grafted and non-budgrafted (tissue culture) plants
  • 4.4 Effect of leaning angle on tension wood formation in mature trees
  • 4.5 Tension wood formation in tapped and untapped zones of the trunk
  • 4.6 Tension wood formation and wind damage
  • 4.7 Visual identification of tension wood in rubber wood logs
  • 4.8 Structural studies on tension wood
  • 4.8.1 Fibres
  • 4.8.1.1 Length of fibres
  • 4.8.1.2 Width of fibres
  • 4.8.1.3 Fibre wall thickness
  • 4.8.2 Vessel elements
  • 4.8.2.1 Length of vessel elements
  • 4.8.2.2 Width of vessel elements
  • 4.8.3 Analysis of pores
  • 4.8.3.1 Number of pores
  • 4.8.3.2 Total area occupied by pores
  • 4.8.3.3 Average area of pores
  • 4.8.4 Analysis of rays
  • 4.8.4.1 Frequency of rays
  • 4.8.4.2 Height of rays
  • 4.8.4.3 Width of rays
  • 4.8.4.4 Height / width ratio of rays
  • 4.9 Histochemical studies
  • 4.9.1 Starch
  • 4.9.2 Cellulose
  • 4.9.3 Lignin
  • 4.9.4 Lipid
  • 4.9.5 Total proteins
  • 4.10 Factors affecting tension wood formation in Hevea
  • 4.10. I Angle of leaning
  • 4.10.1.1 Plants with foliage
  • 4.10.1.2 Plants without foliage
  • 4.10..2 Effect of gravity on tension wood formation
  • 4.10.3 Effect of growth regulators on tension wood formation
  • 4.10.3.1 Apical application of IAA, GA; and TIBA on artificially bent plants
  • 4.10.3.1.1 Application of IAA
  • 4.10.3.1.2 Application of GA;
  • 4.10.3.1.3 Application of TIBA
  • 4.10.3.1.4 Lanolin / Lanolin
  • 4.10.3.2 Lateral application of IAA, GA3 and TIBA on vertically grown plants
  • 4.10.3.2.1 IAA
  • 4.10.3.2.2 GA3
  • 4.10.3.2.3 TIBA
  • 4.10.3.2.4 Lanolin (control)
  • 4.10.3.3 Lateral application of IAA on plants grown vertical through incision of bark
  • 4.10.3.4 Lateral application of IAA and TIBA on artificially bent step- axis at 45
  • 4.10.3.4.1 IAA
  • 4.10.3.4.2 TIBA
  • 10 Cross-sawn discs of rubber wood showing white wooly arcs of compact tension wood (clones-: Tjir 1 & GT 1)
  • 11 Cross-sawn disc of rubber wood showing white wooly arcs of compact tension wood (clones: RRIM 600 & RRII 105)
  • 12 Distribution pattern and structure of tension wood and normal wood (clone Tjir I)
  • 13 Distribution pattern and structure of tension wood and normal wood (clone GT 1)
  • 14 Distribution pattern and structure of tension wood and normal wood (clone RRIM 600)
  • 15 Distribution pattern and structure of tension wood and normal wood (clone RRII 105)
  • 16 Proportion of tension wood in mature trees
  • 17 Cross section of wood showing distribution of tension wood in ten month old budgrafted plants (clones: Tjir I, RRIM 703, PB 5 151 & RRIM 623)
  • 18 Cross section of wood showing distribution of tension wood in ten month old budgrafted plants (clones: GT 1, RRIM 600, PB 217 & PB 235)
  • 19 Cross section of wood showing distribution of tension wood in ten month old budgrafted plants (clones: RRII 105 & GI 1)
  • 20 C.S. of wood showing distribution of tension wood in tissue culture plant and bud grafted plant
  • 21 Tension wood formation and wind damage
  • 22 Macroscopic stainingg of tension wood
  • 23 Morphology of normal and tension wood fibres
  • 24 Length of normal and tension wood fibres from pith to periphery
  • 25 Width of normal wood and tension wood fibers from pith to periphery
  • 26 Length of vessel elements from pith to periphery
  • 27 Width vessel elements from pith to periphery
  • 28 T.L.S. & T.S. of wood showing histochemical localization of starch and cellulose
  • 29 Histochemical localization of lignin in tension wood and normal wood fibres
  • 30 T.L.S. of wood showing localization of lipids T.L.S. of wood showing localization of total proteins
  • 31 C.S of wood showing distribution of tension wood in four month old inclined and vertical seedlings
  • 32 C.S. of wood showing distribution of tension wood in four month old defoliated, inclined and vertical seedlings
  • 33 C.S. of wood showing distribution of tension wood in different segments of the loop
  • 34 C. S. of wood from upper and lower halves of the bent axis treated with IAA and GA3
  • 35 C.S. of wood from upper and lower halves of the bent axis treated with TIBA and lanolin (control)
  • 36 C.S. of wood showing distribution of tension wood in vertical plants treated with lateral application of IAA, GA, and TIBA
  • 37.C.S. of wood showing distribution of tension wood (arrows) in vertical and inclined slants treated with lateral application of IAA and TIBA
  • 5. Discussion
  • 5.1 Distribution pattern of tension wood in Hevea brasiliensis
  • 5.2 Proportion of tension wood in mature trees
  • 5.3 Proportion of tension wood in immature phase
  • 5.4 Proportion of tension wood in bud-grafted and tissue culture plants
  • 5.5 Directional effect of tree leaning on tension wood formation
  • 5.6 Proportion of tension wood in tapped and untapped regions within trees
  • 5.7 Tension wood formation and wind damage
  • 5.8 Visual identification of tension wood
  • 5.9 Structural studies on tension wood
  • 5.9.1 Fibre length
  • 5.9.2 Fibre width
  • 5.9.3 Fibre wall thickness
  • 5.9.4 Length and width of vessel elements
  • 5.9.5 Number of pores
  • 5.9.6 Total area occupied by pores
  • 5.9.7 Average area of pores
  • 5.9.8 Frequency of rays
  • 5.9.9 height. width and height / width ratio of rays
  • 5.10 Histochemical studies
  • 5. 10. l Starch
  • 5.10.2 Lipids
  • 5.10.3 Total Proteins
  • 5.10.4 Cellulose
  • 5.10.5 Lignin
  • 5.11 Factors affecting tension wood formation
  • 5.11.1 Angle of leaning on tension wood formation
  • 5.11.2 Defoliation on tension wood formation
  • 5.11.3 Gravitational response on tension wood formation
  • 5.11.4 Effect of growth regulators on tension wood formation
  • 5.11.4.1 Indole accetic acid (IAA)
  • 5.11.4.2 Gibberellic acid (GA3)
  • 5.11.4.3 Tri-iodo benzoic acid (TIBA)
  • 6. Summary
  • References